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  "author": "Tjitske J. Kooistra (Royal Netherlands Institute for Sea 
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  "author": "Tjitske J. Kooistra (Royal Netherlands Institute for Sea 
            
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Research)",
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Research)",
            
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  "author_email": "tjitske.kooistra@nioz.nl",
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  "author_email": "tjitske.kooistra@nioz.nl",
            
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  "code": "7b.b.ak",
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  "code": "7b.b.ak",
            
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  "contributor": "Royal Netherlands Institute for Sea Research 
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  "contributor": "Royal Netherlands Institute for Sea Research 
            
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(NIOZ)",
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(NIOZ)",
            
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\"Kooistra\",\"orcid\":\"0000-0003-2366-4099\",\"affiliation\":\"Royal 
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\"Kooistra\",\"orcid\":\"0000-0003-2366-4099\",\"affiliation\":\"Royal 
            
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Netherlands Institute for Sea 
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Netherlands Institute for Sea 
            
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actemailaddress\":\"tjitske.kooistra@nioz.nl\",\"formatted\":\"Tjitske 
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actemailaddress\":\"tjitske.kooistra@nioz.nl\",\"formatted\":\"Tjitske 
            
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J. Kooistra - Royal Netherlands Institute for Sea Research - ORCID: 
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J. Kooistra - Royal Netherlands Institute for Sea Research - ORCID: 
            
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0000-0003-2366-4099 - Corresponding Author - 
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0000-0003-2366-4099 - Corresponding Author - 
            
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.kooistra@nioz.nl\"},{\"firstname\":\"Anna-Maartje\",\"lastname\":\"de 
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.kooistra@nioz.nl\"},{\"firstname\":\"Anna-Maartje\",\"lastname\":\"de 
            
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Boer\",\"orcid\":\"0000-0002-7359-6939\",\"affiliation\":\"Wageningen 
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Boer\",\"orcid\":\"0000-0002-7359-6939\",\"affiliation\":\"Wageningen 
            
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University and 
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University and 
            
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uthor\":null,\"contactemailaddress\":null,\"formatted\":\"Anna-Maartje 
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uthor\":null,\"contactemailaddress\":null,\"formatted\":\"Anna-Maartje 
            
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de Boer - Wageningen University and Research - ORCID: 
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de Boer - Wageningen University and Research - ORCID: 
            
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0000-0002-7359-6939\"},{\"firstname\":\"Tjeerd 
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0000-0002-7359-6939\"},{\"firstname\":\"Tjeerd 
            
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\":\"Bouma\",\"orcid\":\"0000-0001-7824-7546\",\"affiliation\":\"Royal 
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\":\"Bouma\",\"orcid\":\"0000-0001-7824-7546\",\"affiliation\":\"Royal 
            
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Netherlands Institute for Sea 
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Netherlands Institute for Sea 
            
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ndingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Tjeerd 
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ndingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Tjeerd 
            
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J. Bouma - Royal Netherlands Institute for Sea Research - ORCID: 
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J. Bouma - Royal Netherlands Institute for Sea Research - ORCID: 
            
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\"Pannozzo\",\"orcid\":\"0000-0002-8082-9251\",\"affiliation\":\"Delft 
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\"Pannozzo\",\"orcid\":\"0000-0002-8082-9251\",\"affiliation\":\"Delft 
            
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University of 
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University of 
            
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ingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Natascia 
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ingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Natascia 
            
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Pannozzo - Delft University of Technology - ORCID: 
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Pannozzo - Delft University of Technology - ORCID: 
            
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0000-0002-8082-9251\"},{\"firstname\":\"Stuart 
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0000-0002-8082-9251\"},{\"firstname\":\"Stuart 
            
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:\"Pearson\",\"orcid\":\"0000-0002-3986-4469\",\"affiliation\":\"Delft 
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:\"Pearson\",\"orcid\":\"0000-0002-3986-4469\",\"affiliation\":\"Delft 
            
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University of 
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University of 
            
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ndingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Stuart 
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ndingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Stuart 
            
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G. Pearson - Delft University of Technology - ORCID: 
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G. Pearson - Delft University of Technology - ORCID: 
            
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0000-0002-3986-4469\"},{\"firstname\":\"Ad\",\"lastname\":\"van der 
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0000-0002-3986-4469\"},{\"firstname\":\"Ad\",\"lastname\":\"van der 
            
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espondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Ad 
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espondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Ad 
            
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van der Spek - Deltares - ORCID: 
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van der Spek - Deltares - ORCID: 
            
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0000-0001-7131-4363\"},{\"firstname\":\"Henko\",\"lastname\":\"de 
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0000-0001-7131-4363\"},{\"firstname\":\"Henko\",\"lastname\":\"de 
            
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Stigter\",\"orcid\":\"0000-0002-4878-6546\",\"affiliation\":\"Royal 
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Stigter\",\"orcid\":\"0000-0002-4878-6546\",\"affiliation\":\"Royal 
            
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Netherlands Institute for Sea 
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Netherlands Institute for Sea 
            
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ondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Henko 
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ondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Henko 
            
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de Stigter - Royal Netherlands Institute for Sea Research - ORCID: 
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de Stigter - Royal Netherlands Institute for Sea Research - ORCID: 
            
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linga\",\"orcid\":\"0000-0003-4061-3066\",\"affiliation\":\"Wageningen 
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linga\",\"orcid\":\"0000-0003-4061-3066\",\"affiliation\":\"Wageningen 
            
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University and 
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University and 
            
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ondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Jakob 
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ondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Jakob 
            
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Wallinga - Wageningen University and Research - ORCID: 
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Wallinga - Wageningen University and Research - ORCID: 
            
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\"Witbaard\",\"orcid\":\"0000-0002-7062-7920\",\"affiliation\":\"Royal 
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\"Witbaard\",\"orcid\":\"0000-0002-7062-7920\",\"affiliation\":\"Royal 
            
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Netherlands Institute for Sea 
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Netherlands Institute for Sea 
            
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spondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Rob 
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spondingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Rob 
            
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Witbaard - Royal Netherlands Institute for Sea Research - ORCID: 
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Witbaard - Royal Netherlands Institute for Sea Research - ORCID: 
            
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\"Soetaert\",\"orcid\":\"0000-0003-4603-7100\",\"affiliation\":\"Royal 
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\"Soetaert\",\"orcid\":\"0000-0003-4603-7100\",\"affiliation\":\"Royal 
            
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Netherlands Institute for Sea 
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Netherlands Institute for Sea 
            
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dingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Karline 
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dingauthor\":null,\"contactemailaddress\":null,\"formatted\":\"Karline 
            
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Soetaert - Royal Netherlands Institute for Sea Research - ORCID: 
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Soetaert - Royal Netherlands Institute for Sea Research - ORCID: 
            
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  "deposit_date": "2025-12-01",
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  "deposit_date": "2025-12-01",
            
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  "depositor": "Tjitske Kooistra (tjitske.kooistra@nioz.nl)",
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  "depositor": "Tjitske Kooistra (tjitske.kooistra@nioz.nl)",
            
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  "distribution_date": "2025-12-01",
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  "distribution_date": "2025-12-01",
            
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  "distributor": "Research Data Management(NIOZ Royal Netherlands 
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  "distributor": "Research Data Management(NIOZ Royal Netherlands 
            
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Institute for Sea Research)",
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Institute for Sea Research)",
            
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  "doi_date_published": "2026-04-14T10:25:32.963757",
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  "maintainer": "Tjitske",
            
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  "metadata_modified": "2026-04-14T08:26:03.403040",
            
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  "name": 
73
  "name": 
            
74
on-bioturbation-and-physical-mixing-on-an-intertidal-flat-by-combini",
74
on-bioturbation-and-physical-mixing-on-an-intertidal-flat-by-combini",
            
75
  "notes": "Sediment transport and seabed composition can both be 
75
  "notes": "Sediment transport and seabed composition can both be 
            
76
influenced by bioturbation and hydrodynamically driven sediment mixing 
76
influenced by bioturbation and hydrodynamically driven sediment mixing 
            
77
and deposition. In a dynamic intertidal environment, it is challenging 
77
and deposition. In a dynamic intertidal environment, it is challenging 
            
78
to distinguish the relative contribution of both processes. We aim to 
78
to distinguish the relative contribution of both processes. We aim to 
            
79
unravel their relative importance by combining several tracers, each 
79
unravel their relative importance by combining several tracers, each 
            
80
having its own specific timescale and target particle size. We 
80
having its own specific timescale and target particle size. We 
            
81
combined (1) 210Pb that quantifies long-term (years\u2013decades) 
81
combined (1) 210Pb that quantifies long-term (years\u2013decades) 
            
82
mixing of fine sediment fractions with (2) Chlorophyll-a and (3) 
82
mixing of fine sediment fractions with (2) Chlorophyll-a and (3) 
            
83
luminophores that both quantify short-term mixing of fine sediment 
83
luminophores that both quantify short-term mixing of fine sediment 
            
84
fractions (days\u2013weeks), and (4) multi-grain quartz and 
84
fractions (days\u2013weeks), and (4) multi-grain quartz and 
            
85
single-grain feldspar luminescence dating, which use the bleaching of 
85
single-grain feldspar luminescence dating, which use the bleaching of 
            
86
sand grains\u2019 inherent luminescence signal by light to assess 
86
sand grains\u2019 inherent luminescence signal by light to assess 
            
87
mixing of sand and thereby quantifies long-term mixing. Single grain 
87
mixing of sand and thereby quantifies long-term mixing. Single grain 
            
88
feldspar luminescence is here for the first time applied in the 
88
feldspar luminescence is here for the first time applied in the 
            
89
intertidal environment. We compare results for a sandy and a muddy 
89
intertidal environment. We compare results for a sandy and a muddy 
            
90
intertidal flat at the island of Texel (Wadden Sea, the Netherlands), 
90
intertidal flat at the island of Texel (Wadden Sea, the Netherlands), 
            
91
each with their own characteristic benthic community. Recent 
91
each with their own characteristic benthic community. Recent 
            
92
bioturbation became apparent from Chlorophyll-a and luminophore 
92
bioturbation became apparent from Chlorophyll-a and luminophore 
            
93
profiles: particles were rapidly reworked to a depth of decimetres. 
93
profiles: particles were rapidly reworked to a depth of decimetres. 
            
94
210Pb also suggested mixing and non-local exchange of particles by 
94
210Pb also suggested mixing and non-local exchange of particles by 
            
95
bioturbation. The combination of luminescence signals suggested that 
95
bioturbation. The combination of luminescence signals suggested that 
            
96
after deposition, not all sand grains did resurface repeatedly and for 
96
after deposition, not all sand grains did resurface repeatedly and for 
            
97
longer time periods through bioturbation. Coarse- and fine-grained 
97
longer time periods through bioturbation. Coarse- and fine-grained 
            
98
tracer profiles show the differential behaviour and reworking of the 
98
tracer profiles show the differential behaviour and reworking of the 
            
99
mud and sand fraction within the sediment matrix: as expected with 
99
mud and sand fraction within the sediment matrix: as expected with 
            
100
particle-selective bioturbation, mud is preferentially bioturbated and 
100
particle-selective bioturbation, mud is preferentially bioturbated and 
            
101
infiltrates passively, while sand grains have a higher ability to 
101
infiltrates passively, while sand grains have a higher ability to 
            
102
conserve layering. Single-grain feldspar luminescence is a promising 
102
conserve layering. Single-grain feldspar luminescence is a promising 
            
103
technique to demonstrate the long-term reworking of sand grains, 
103
technique to demonstrate the long-term reworking of sand grains, 
            
104
however, in young and dynamic environments, a combination of tracers 
104
however, in young and dynamic environments, a combination of tracers 
            
105
remains necessary to inform on the origin of mixing.",
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remains necessary to inform on the origin of mixing.",
            
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